A set of methods and equations was developed in order to predict total and component biomass and volume for trees in the Forest Inventory and Analysis Database (FIADB). The goals/requirements were to:
Data were compiled by several university collaborators and the Legacy Tree Data website. The fitting dataset consisted of 234,823 destructively sampled trees from 339 species across 23 ecological divisions. Four candidate allometric models were selected for evaluation:
\(Y = a * D^b * H^c\) (Schumacher-Hall)
\(\begin{aligned} Y = \begin{cases} & a_0 * D^{b_0} * H^c; D < k \\ & a_0 * k^{(b_0 - b_1)} * D^b_1 * H^c; D \ge k \end{cases} \end{aligned}\) (Segmented) | where \(k=9\) for softwood trees and \(k=11\) for hardwoods
\(Y = a * D^{(a_1 * (1 - e^{(-b_1 * D)})^{c_1})} * H^c\) (Continuously Variable)
\(Y = a * D^b * H^c * e^{(-(b_2 * D))}\) (Modified Wiley)
All candidate models were evaluated for each species. The Schumacher-Hall model was considered the ‘default’ equation form. In order for a different equation to be chosen, it needed to have a lower AIC score and all estimated coefficients needed to be significant at the \(\alpha = 0.05\) level.
Preliminary work showed that the relationship between tree size and volume/biomass of a species frequently varied across ecological divisions (Figure 1). Therefore, models were fit by species within ecological division (Figure 2). Within-division biomass models (total, stem wood, stem bark, branch, foliage) were developed for any groups with at least fifty observations. Within-division volume models (stem wood, stem bark, volume ratio) were developed for groups with at least eighty observations. As FIADB contained species within division combinations that were not represented in the fitting dataset, species-level models were also fit across divisions.
The species-level models, either within division or across divisions, accounted for 89% of standing volume in FIADB and 72% of standing aboveground biomass. In order to produce estimates for the remaining species in FIADB, models were also estimated for the Jenkins species groups. The Jenkins groups are collections of species created according to phylogenetic relationships and wood specific gravity, and are already in use by FIA. Models were estimated for eight of the ten Jenkins groups (Douglas-fir and woodland groups were excluded due to the former group being a single species and lack of data for the latter). For species with between five and fifty/eighty observations, mixed-effects models were estimated at the Jenkins group level that used species as a random effect. For species with fewer than five observations, a modified version of the Schumacher-Hall that incorporates published species-level wood density values was estimated at the Jenkins group.
Allometric models were developed for: 1. total stem wood volume, 2. total stem bark volume, 3. total branch wood and bark weight, 4. total aboveground biomass (without foliage), and 5. total foliage biomass
Additionally, inside- and outside-bark volume ratio models were estimated for all possible species and Jenkins groups:
These ratio equations could were needed to find the proportion of volume between two heights, as well as estimate the height to any diameter on the stem.
The following illustrates how the prediction system functions for all trees in FIADB:
Steps 12 & 13 ensure that the main stem can be broken into any sub-components (i.e., stump, merchantable bole, top) and still be additive with the adjusted total stem weight. Using the same set of ratio-equation coefficients (step 4) insures that all outside-bark volumes are larger than the inside-bark volumes
In order to examine the potential changes in estimated volume and biomass using the new system, the updated modeling algorithm was applied to FIADB. The 2017 evaluation was used for all states (except Hawaii) and the models were applied to all trees except for woodland species and a small number of saplings in the Northeast without measured total heights. Cull deductions were applied in the same way as currently used. Population-level estimates using the new system were produced for six components (merchantable stem wood volume, sawlog wood volume, merchantable bole biomass, top and limb biomass, stump weight, and total aboveground biomass), and were compared to current estimates.
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